diff --git a/modules/archive/lib.py b/modules/archive/lib.py
index 2c271228..ba90469e 100644
--- a/modules/archive/lib.py
+++ b/modules/archive/lib.py
@@ -1018,10 +1018,12 @@ def domain_download_links(session: Session) -> Dict[int, int]:
 
     links = dict()
     downloads = session.query(Download.id, Download.url, Download.downloader, Download.sub_downloader,
-                              Download.settings) \
+                              Download.destination, Download.settings) \
         .filter(Download.frequency.isnot(None), Download.collection_id.is_(None))
-    for id_, url, downloader, sub_downloader, settings in downloads:
-        if destination := (settings or dict()).get('destination'):
+    for id_, url, downloader, sub_downloader, destination, settings in downloads:
+        # An old Download kept its destination in its settings.
+        destination = str(destination) if destination else (settings or dict()).get('destination')
+        if destination:
             for collection_id, directory, prefix in directories:
                 if destination == directory or destination.startswith(prefix):
                     links[id_] = collection_id
diff --git a/modules/archive/test/test_lib.py b/modules/archive/test/test_lib.py
index 316f58b1..2d6379e7 100644
--- a/modules/archive/test/test_lib.py
+++ b/modules/archive/test/test_lib.py
@@ -1277,6 +1277,30 @@ def test_link_domain_and_downloads(test_session, test_download_manager):
     assert download4.collection_id is None  # no frequency = no link
 
 
+def test_link_domain_and_downloads_by_destination(test_session, test_directory, test_download_manager):
+    """A Download whose destination column is in a domain's directory is linked to that domain (the destination was
+    moved out of the settings)."""
+    from wrolpi.downloader import Download
+    from modules.archive.lib import link_domain_and_downloads
+
+    collection = Collection(name='example.com', kind='domain', directory=test_directory / 'archive/example.com')
+    test_session.add(collection)
+    exact = Download(url='https://other.com/rss', downloader='rss', sub_downloader='archive', frequency=86400,
+                     destination=test_directory / 'archive/example.com')
+    below = Download(url='https://other.com/rss2', downloader='rss', sub_downloader='archive', frequency=86400,
+                     destination=test_directory / 'archive/example.com/2025/01')
+    similar = Download(url='https://other.com/rss3', downloader='rss', sub_downloader='archive', frequency=86400,
+                       destination=test_directory / 'archive/example.com.au')
+    test_session.add_all([exact, below, similar])
+    test_session.commit()
+
+    link_domain_and_downloads(test_session)
+
+    assert exact.collection_id == collection.id
+    assert below.collection_id == collection.id
+    assert similar.collection_id is None
+
+
 @skip_ci
 @pytest.mark.asyncio
 async def test_archive_download_uses_domain_collection_directory(async_client, test_session, test_directory, fake_now,